A cactus (plural: cacti or cactuses) is a member of the plant Plants are living organisms belonging to the kingdom Plantae. They include familiar organisms such as trees, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The scientific study of plants, known as botany, has identified about 350,000 extant species of plants, defined as seed plants, bryophytes, ferns and fern allies. As of 2004, family Cactaceae, native to the Americas The Americas, or America, are lands in the Western hemisphere, also known as the New World, comprising the continents of North America and South America with their associated islands and regions. The plural form the Americas is often used in English, as the singular America is ambiguous: America is more commonly used to refer to the United States (with one exception, Rhipsalis baccifera, which is native to parts of the Old World The Old World consists of those parts of Earth known to Europeans[note], Asians and Africans in the 15th century. It is used in the context of, and contrast with, the "New World"). They are often used as ornamental plants Ornamental plants or garden plants are typically grown in the flower garden or as house plants. Most commonly they are grown for the display of their flowers. Other common ornamental features include leaves, scent, fruit, stem and bark. In some cases, unusual features may be considered ornamental, such as the prominent and rather vicious thorns of, and some are also crop A crop is the annual or season's yield of any plant that is grown in significant quantities to be harvested as food, as livestock fodder, fuel, or for any other economic purpose. This category includes crop species as well as agricultural techniques related to cropping plants for fodder, forage, fruits, cochineal The Cochineal is a scale insect in the suborder Sternorrhyncha, from which the crimson-coloured dye carmine is derived. A primarily sessile parasite native to tropical and subtropical South America and Mexico, this insect lives on cacti from the genus Opuntia, feeding on plant moisture and nutrients, and other uses. Numerous species have been used since ancient times by indigenous peoples for their psychedelic The term psychedelic is derived from the Greek words ψυχή and δηλοῦν (deloun, "to manifest"), translating to "mind-manifesting". A psychedelic experience is characterized by the perception of aspects of one's mind previously unknown, or by the creative exuberance of the mind liberated from its ostensibly ordinary effects. Cacti are part of the plant order Caryophyllales, which also includes members like beets The beet is a plant in the Chenopodiaceae family. It is best known in its numerous cultivated varieties, the most well known of which is probably the red or purple root vegetable known as the beetroot or garden beet. However, other cultivated varieties include the leaf vegetables chard and spinach beet, as well as the root vegetables sugar beet,, gypsophila, spinach Spinach is an edible flowering plant in the family of Amaranthaceae. It is native to central and southwestern Asia. It is an annual plant (rarely biennial), which grows to a height of up to 30 cm. Spinach may survive over winter in temperate regions. The leaves are alternate, simple, ovate to triangular-based, very variable in size from about 2-30, amaranth Amaranthus, collectively known as amaranth, is a cosmopolitan genus of herbs. Approximately 60 species are recognized, with inflorescences and foliage ranging from purple and red to gold. Members of this genus share many characteristics and uses with members of the closely related genus Celosia, tumbleweeds A tumbleweed is the above-ground part of a plant that, once mature and dry, separates from the root and tumbles away in the wind. Usually, the tumbleweed is the entire plant apart from the roots, but in a few species it is a flower cluster. The tumbleweed habit is most common in steppe and desert plants. The tumbleweed is a diaspore, aiding in, carnations Dianthus caryophyllus is a species of Dianthus. It is probably native to the Mediterranean region but its exact range is unknown due to extensive cultivation for the last 2,000 years. It is the wild ancestor of the garden Carnation, rhubarb They are herbaceous perennial plants growing from short, thick rhizomes. They have large leaves that are somewhat triangular shaped with long fleshy petioles. They have small flowers grouped in large compound leafy greenish-white to rose-red inflorescences, buckwheat Buckwheat refers to a variety of plants in the dicot family Polygonaceae: the Eurasian genus Fagopyrum, the North American genus Eriogonum, and the Northern Hemisphere genus Fallopia. Either of the latter two may be referred to as wild buckwheat, plumbago, bougainvillea Bougainvillea is a genus of flowering plants native to South America from Brazil west to Peru and south to southern Argentina (Chubut Province). Different authors accept between four and 18 species in the genus. The plant was classified by Europeans in Brazil in 1768, by Philibert Commerçon, a French botanist accompanying French Navy admiral and, chickweed and knotgrass.

Cacti are unusual and distinctive plants Plants are living organisms belonging to the kingdom Plantae. They include familiar organisms such as trees, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The scientific study of plants, known as botany, has identified about 350,000 extant species of plants, defined as seed plants, bryophytes, ferns and fern allies. As of 2004,, which are adapted to extremely arid A region is said to be arid when it is characterized by a severe lack of available water, to the extent of hindering or even preventing the growth and development of plant and animal life. As a result, environments subject to arid climates tend to lack vegetation and are called xeric or desertic and/or semi-arid A semi-arid climate or steppe climate describes climatic regions that receive precipitation below potential evapotranspiration, but not extremely so. A more precise definition is given by the Köppen climate classification that treats steppe climates as intermediates between desert climates (BW) and humid climates in ecological characteristics and hot environments The biophysical environment is the symbiosis between the physical environment and the biological life forms within the environment, and includes all variables that comprise the Earth's biosphere. The biophysical environment can be divided into two categories: the natural environment and the built environment, with some overlap between the two, as well as tropical environments as epiphytes Epiphyte is one of the subdivisions of the Raunkiær system. The term most commonly refers to higher plants, but epiphytic bacteria, fungi , algae, lichens, mosses, and ferns exist as well. The term epiphytic derives from the Greek epi- (meaning 'upon') and phyton (meaning 'plant'). Epiphytic plants are sometimes called "air plants" or hemi-epiphytes A hemiepiphyte is a plant which begins its life as an epiphyte but which later grows roots down into the ground. The seeds of hemiepiphytes germinate in the canopy and initially live epiphytically. They send roots downward, and these roots eventually make contact with the ground.[1][2][3] They show a wide range of anatomical Anatomy is a branch of biology and medicine that is the consideration of the structure of living things. It is a general term that includes human anatomy, animal anatomy (zootomy) and plant anatomy (phytotomy). In some of its facets anatomy is closely related to embryology, comparative anatomy and comparative embryology, through common roots in and physiological Physiology is the science of the functioning of living systems. It is a subcategory of biology. In physiology, the scientific method is applied to determine how organisms, organ systems, organs, cells and biomolecules carry out the chemical or physical function that they have in a living system. The word physiology is from Ancient Greek: φύσις features which conserve water. Their stems have adapted to become photosynthetic Photosynthesis is a process that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight. Photosynthesis occurs in plants, algae, and many species of bacteria, but not in archaea. Photosynthetic organisms are called photoautotrophs, since they can create their own food. In plants, algae, and cyanobacteria, and succulent Succulent plants, also known as succulents or fat plants, are water-retaining plants adapted to arid climate or soil conditions. Succulent plants store water in their leaves, stems, and also in roots. Geophytes that survive unfavourable periods by dying back to underground storage organs such as tuberous roots, corms, bulbs, and rhizomes, may be, while the leaves have become the spines for which cacti are well known.

Cacti come in a wide range of shapes and sizes. The tallest is Pachycereus pringlei, with a maximum recorded height of 19.2 m,[4] and the smallest is Blossfeldia liliputiana, only about 1 cm diameter at maturity.[5] Cactus flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to mediate the union of male sperm with female ovum in order to produce seeds. The process begins with pollination, is followed by fertilization, leading to the formation and dispersal of the seeds. For are large, and like the spines and branches arise from areoles Areoles are the distinctive feature of cacti, and identify them as a separate family from other succulent plants. Areoles give rise to spines or, on certain cacti, small, detachable glochids which are an additional form of protection. The areoles on cacti are clearly visible. They generally appear as small light to dark colored bumps, out of which.

Closeup image of a cactus flower (Echinopsis spachiana) showing large number of stamens. Closeup look of a cactus perianth.

Cacti are perennial and grow as trees, shrubs, or vines. Most species are terrestrial, but there are also many epiphytic Epiphyte is one of the subdivisions of the Raunkiær system. The term most commonly refers to higher plants, but epiphytic bacteria, fungi , algae, lichens, mosses, and ferns exist as well. The term epiphytic derives from the Greek epi- (meaning 'upon') and phyton (meaning 'plant'). Epiphytic plants are sometimes called "air plants" species, especially in the tribes Rhipsalideae and Hylocereeae. In most species, except for the sub-family Pereskioideae (see image), the leaves are greatly or entirely reduced. The leaves may also be tiny and deciduous Deciduous means falling off at maturity or tending to fall off and is typically used in reference to trees or shrubs that lose their leaves seasonally and to the shedding of other plant structures such as petals after flowering or fruit when ripe. In a more specific sense deciduous means the dropping of a part that is no longer needed, or falling as can be seen on new shoots of Opuntia Opuntia, also known as nopales or paddle cactus , is a genus in the cactus family, Cactaceae. Spines grown by the cacti are actually modified leaves; the stems (the green "pads" of many cacti) have also evolved to photosynthesize. The study of when spines grow and how they can be used to tell the cactus' age is called acanthochronology. The flowers, mostly radially symmetrical and bisexual, bloom either by day or by night, depending on the species. Their shape varies from tube-like through bell-like to wheel-shaped, and their size from 0.2 to 15–30 centimetres. Most of them have numerous sepals A sepal is a part of the flower of angiosperms (flowering plants). Sepals in most flowers are green and lie under the more conspicuous petals. However, the term tepal is usually applied when the petals and sepals share the same color, or the petals are absent and the sepals are colorful. When the flower is in bud, the sepals enclose and protect (from 5 to 50 or more), and change form from outside to inside, from bracts In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis, or cone scale. Bracts are often different from foliage leaves, for example being smaller, larger, or of a different color or texture to petals A petal is one member or part of the corolla of a flower. The corolla is the name for all of the petals of a flower. The corolla (petals) and calyx (the name for all the sepals) make up the outer whorls of a flower; these form what is known as the perianth, which attracts pollinators with its bright color. The term "tepal" is usually. They have stamens The stamen is the male reproductive organ of a flower. Each stamen generally has a stalk called the filament (from Latin filum, meaning "thread"), and, on top of the filament, an anther (from Ancient Greek anthera, feminine of antheros "flowery," from anthos "flower"), and pollen sacs, called microsporangia. The in great numbers (from 50 to 1,500, rarely fewer). Nearly all species of cacti have a bitter mucilaginous sap contained within them. The berry-like fruits may contain few to many (3,000), seeds, which can be between 0.4 and 12 mm long.[6]

The life of a cactus is seldom longer than 300 years[citation needed], but may be as short as 25 years, (although these flower as early as their second year). The Saguaro cactus (Carnegiea gigantea) grows to a height of up to 15 metres (the record is 17 metres 67 cm), but in its first ten years, it grows only 10 centimeters. The "mother-in-law's cushion" (Echinocactus grusonii) reaches a height of 2.5 meters and a diameter of 1 metre and – at least on the Canaries The Canary Islands (pronounced /kəˈnɛəriː ˈaɪləndz/, colloquially also known as the Canaries; Spanish: Islas Canarias, pronounced [ˈislas kaˈnarjas]; 28°06′N 15°24′W / 28.1°N 15.4°W – is already capable of flowering after 6 years. The diameter of cactus flowers ranges from 5 to 30 cm; the colors are often conspicuous and spectacular.

The cactus family is endemic to the Americas The Americas, or America, are lands in the Western hemisphere, also known as the New World, comprising the continents of North America and South America with their associated islands and regions. The plural form the Americas is often used in English, as the singular America is ambiguous: America is more commonly used to refer to the United States with one exception, Rhipsalis baccifera; this species has a pantropical In biology, pantropical means that the organism occurs in the tropical regions of all of the major continents, i.e. in Africa, in Asia and in the Americas distribution, occurring in the Old World The Old World consists of those parts of Earth known to Europeans[note], Asians and Africans in the 15th century. It is used in the context of, and contrast with, the "New World" tropical Africa Africa is the world's second-largest and second most-populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area. With a billion people (as of 2009, see table) in 61 territories, it accounts for about 14.72% of the world's human population, Madagascar Madagascar, or Republic of Madagascar , is an island nation in the Indian Ocean off the southeastern coast of Africa. The main island, also called Madagascar, is the fourth-largest island in the world and Sri Lanka As a result of its location in the path of major sea routes, Sri Lanka is a strategic naval link between West Asia and South East Asia.[citation needed] It has also been a center of the Buddhist religion and culture from ancient times and is one of the few remaining abodes of Buddhism in South Asia, including Ladakh, Bhutan and the Chittagong Hill as well as in tropical America. This plant is thought to be a relatively recent colonist in the Old World The Old World consists of those parts of Earth known to Europeans[note], Asians and Africans in the 15th century. It is used in the context of, and contrast with, the "New World" (within the last few thousand years), probably carried as seeds A seed ( /ˈsiːd/ ) is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant. The formation of the seed completes the process of reproduction in seed in the digestive tracts The Human gastrointestinal tract is the system by which ingested food is acted upon by physical and chemical means to provide the body with nutrients it can absorb and to excrete waste products; in mammals the system includes the alimentary canal extending from the mouth to the anus, and the hormones and enzymes assisting in digestion of migratory Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. These however are usually irregular or in only one direction and are termed variously as nomadism, invasions, dispersal or irruptions. Migration is marked by its birds Birds are winged, bipedal, endothermic (warm-blooded), egg-laying, vertebrate animals. There are around 10,000 living species, making them the most varied of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from the 5 cm (2 in) Bee Hummingbird to the 2.75 m (9 ft) Ostrich. Many other cacti have become naturalized to similar environments in other parts of the world after being introduced by people, e.g. Australia For at least 40,000 years before European settlement in the late 18th century, Australia was inhabited by indigenous Australians, who belonged to one or more of the roughly 250 language groups. After sporadic visits by fishermen from the immediate north and discovery by Dutch explorers in 1606, Australia's eastern half was claimed by Britain in 177, Hawaii The state encompasses nearly the entire volcanic Hawaiian Island chain, which comprises hundreds of islands spread over 1,500 miles . At the southeastern end of the archipelago, the eight "main islands" are (from the northwest to southeast) Niʻihau, Kauaʻi, Oʻahu, Molokaʻi, Lānaʻi, Kahoʻolawe, Maui, and Hawaiʻi. The last is by, and the Mediterranean region The Mediterranean Basin comprises the lands around and surrounded by the Mediterranean Sea. In biogeography, the Mediterranean Basin refers to the lands around the Mediterranean Sea that have a Mediterranean climate, with mild, rainy winters and hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation. The Tehuacán Valley of Mexico In Pre-Columbian Mesoamerica many cultures matured into advanced civilizations such as the Olmec, the Toltec, the Teotihuacan, the Zapotec, the Maya and the Aztec before the first contact with Europeans. In 1521, Spain conquered and colonized the territory, which was administered as the viceroyalty of New Spain which would eventually become Mexico has one of the richest occurrences of cacti in the world.[7] Species diversity decreases as one travels north; hundreds of species can be found in areas of Mexico, dozens of species are found in the Sonora and Mojave deserts of the southwestern U.S., and only several species are found in the eastern plains and dry valleys of Montana and Alberta.

Cacti are believed to have evolved in the last 30 to 40 million years[citation needed]. Long ago, the Americas were joined to the other continents, but separated due to continental drift. Unique species in the New World must have developed after the continents had moved apart or began developing just prior to the separation. Significant distance between the continents was only achieved in around the last 50 million years. This may explain why cacti are so rare in Africa as the continents had already separated when cacti evolved. Many succulent plants in both the Old and New World bear a striking resemblance to cacti, and are often called "cactus" in common usage.

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