Anthophyta Magnoliophyta Cronquist Arthur John Cronquist was a North American botanist and a specialist on Compositae. He is considered one of the most influential botanists of the 20th century, largely due to his formulation of the Cronquist system. Two plant genera in the aster family have been named in his honor. These are Cronquistia, a possible synonym of Carphochaete, and, Takht. Armen Leonovich Takhtajan or Takhtajian (June 10, 1910 – November 13, 2009), was a Soviet-Armenian botanist, one of the most important figures in 20th century plant evolution and systematics and biogeography. His other interests included morphology of flowering plants, paleobotany, and the flora of the Caucasus. He was born in Shushi & W.Zimm. Walter Max Zimmerman was a German botanist. The standard botanical author abbreviation W.Zimm. is applied to species he described, 1966
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The flowering plants (angiosperms), also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants The embryophytes are the most familiar group of plants. They are often called land plants because they live primarily in terrestrial habitats, in contrast with the related green algae that are primarily aquatic. The embryophytes include trees, flowers, ferns, mosses, and various other green land plants. All are complex multicellular eukaryotes. Together with gymnosperms Once an authoritative and major classification level in the plant kingdom, the gymnosperms form today a rather heterogeneous group of seed-bearing plants that includes conifers, cycads, Ginkgo and Gnetales. The term "gymnosperm" comes from the Greek word gymnospermos , meaning "naked seeds", after the unenclosed condition of, they are the only extant groups of seed-producing plants The spermatophytes (also known as phanerogams) comprise those plants that produce seeds. They are a subset of the embryophytes or land plants. The living spermatophytes form five groups:, but they can be distinguished from the gymnosperms by a series of synapomorphies In cladistics, a synapomorphy or synapomorphic character is a trait that is shared by two or more taxa and their most recent common ancestor, whose ancestor in turn does not possess the trait. A synapomorphy is thus an apomorphy visible in multiple taxa, where the trait in question originates in their last common ancestor. The word " (derived characteristics A trait is a distinct variant of a phenotypic character of an organism that may be inherited, environmentally determined or somewhere in between. For example, eye color is a character or abstraction of an attribute, while blue, brown and hazel are traits). These characteristics include flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to mediate the union of male sperm with female ovum in order to produce seeds. The process begins with pollination, is followed by fertilization, leading to the formation and dispersal of the seeds. For, endosperm Endosperm is the tissue produced under the seeds of most flowering plants around the time of fertilization. It surrounds the embryo and provides nutrition in the form of starch, though it can also contain oils and protein. This makes endosperm an important source of nutrition in human diet. For example, wheat endosperm is ground into flour for within the seeds, and the production of fruits The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state, such as apples, oranges, grapes, strawberries, juniper berries and bananas. Seed-associated structures that do not fit these that contain the seeds.
The ancestors of flowering plants diverged from gymnosperms Once an authoritative and major classification level in the plant kingdom, the gymnosperms form today a rather heterogeneous group of seed-bearing plants that includes conifers, cycads, Ginkgo and Gnetales. The term "gymnosperm" comes from the Greek word gymnospermos , meaning "naked seeds", after the unenclosed condition of around 245–202 million years ago, and the first flowering plants known to exist are from 140 million years ago. They diversified enormously during the Lower Cretaceous and became widespread around 100 million years ago, but replaced conifers The conifers, division Pinophyta, also known as division Coniferophyta or Coniferae, are one of 13 or 14 division level taxa within the Kingdom Plantae. Pinophytes are gymnosperms. They are cone-bearing seed plants with vascular tissue; all extant conifers are woody plants, the great majority being trees with just a few being shrubs. Typical as the dominant trees only around 60-100 million years ago.
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Angiosperm derived characteristics
The flowers, which are the reproductive organs A sex organ, or primary sexual characteristic, as narrowly defined, is any of the anatomical parts of the body which are involved in sexual reproduction and constitute the reproductive system in a complex organism; flowers are the reproductive organs of flowering plants, cones are the reproductive organs of coniferous plants, whereas mosses, ferns, of flowering plants, are the most remarkable feature distinguishing them from other seed plants. Flowers aid angiosperms by enabling a wider range of adaptability and broadening the ecological niches In ecology, a niche is a term describing the relational position of a species or population in its ecosystem to each other; e.g. a dolphin could potentially be in another ecological niche from one that travels in a different pod if the members of these pods utilize significantly different food resources and foraging methods. A shorthand definition open to them. This has allowed flowering plants to largely dominate terrestrial An ecoregion , sometimes called a bioregion, is an ecologically and geographically defined area that is smaller than an ecozone and larger than an ecosystem. Ecoregions cover relatively large areas of land or water, and contain characteristic, geographically distinct assemblages of natural communities and species. The biodiversity of flora, fauna ecosystems.
Stamens are much lighter than the corresponding organs of gymnosperms and have contributed to the diversification of angiosperms through time with adaptations Adaptation is the evolutionary process whereby a population becomes better suited to its habitat. This process takes place over many generations, and is one of the basic phenomena of biology to specialized pollination Pollination is the process by which pollen is transferred in plants, thereby enabling fertilization and sexual reproduction. Pollen grains, which contain the male gametes to where the female gamete(s) are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself. The receptive part of the carpel is called a syndromes, such as particular pollinators. Stamens have also become modified through time to prevent self-fertilization, which has permitted further diversification, allowing angiosperms eventually to fill more niches.
- Reduced male parts, three cells
The male gametophyte A gametophyte is the haploid, multicellular phase of plants and algae that undergo alternation of generations, with each of its cells containing only a single set of chromosomes in angiosperms is significantly reduced in size compared to those of gymnosperm seed plants. The smaller pollen decreases the time from pollination — the pollen grain reaching the female plant — to fertilization Fertilisation , is the fusion of gametes to produce a new organism. In animals, the process involves the fusion of an ovum with a sperm, which eventually leads to the development of an embryo. Depending on the animal species, the process can occur within the body of the female in internal fertilisation, or outside in the case of external of the ovary; in gymnosperms fertilization can occur up to a year after pollination, while in angiosperms the fertilization begins very soon after pollination. The shorter time leads to angiosperm plants setting seeds sooner and faster than gymnosperms, which is a distinct evolutionary advantage.
- Closed carpel The gynoecium is a term with several meanings in botanical usage. In reference to mosses; liverworts; and hornworts, gynoecium refers to a cluster of archegonia and any associated modified leaves or stems present on a gametophyte shoot. The more common use of gynoecium, however, is to refer to the ovule-producing part of a flower. The gynoecium in enclosing the ovules (carpel or carpels and accessory parts may become the fruit The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state, such as apples, oranges, grapes, strawberries, juniper berries and bananas. Seed-associated structures that do not fit these)
The closed carpel of angiosperms also allows adaptations to specialized pollination syndromes and controls. This helps to prevent self-fertilization, thereby maintaining increased diversity. Once the ovary is fertilized, the carpel and some surrounding tissues develop into a fruit. This fruit often serves as an attractant to seed-dispersing animals. The resulting cooperative relationship presents another advantage to angiosperms in the process of dispersal Biological dispersal refers to a species movement away from an existing population or away from the parent organism. Through simply moving from one habitat patch to another, the dispersal of an individual has consequences not only for individual fitness, but also for population dynamics, population genetics, and species distribution. Understanding.
- Reduced female gametophyte, seven cells with eight nuclei
The reduced female gametophyte, like the reduced male gametophyte, may be an adaptation allowing for more rapid seed set, eventually leading to such flowering plant adaptations as annual herbaceous life cycles, allowing the flowering plants to fill even more niches.
Endosperm formation generally begins after fertilization and before the first division of the zygote A zygote , or zygocyte, is the initial cell formed when a new organism is produced by means of sexual reproduction. A zygote is synthesized from the union of two gametes, and constitutes the first stage in a unique organism's development. Zygotes are usually produced by a fertilization event between two haploid cells — an ovum from a female and. Endosperm is a highly nutritive tissue that can provide food for the developing embryo An embryo is a multicellular diploid eukaryote in its earliest stage of development, from the time of first cell division until birth, hatching, or germination. In humans, it is called an embryo until about eight weeks after fertilization (i.e. ten weeks LMP), and from then it is instead called a fetus, the cotyledons, and sometimes for the seedling A seedling is a young plant sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle , the hypocotyl (embryonic shoot), and the cotyledons (seed leaves). The two classes of flowering plants are distinguished by their numbers when it first appears.
These distinguishing characteristics taken together have made the angiosperms the most diverse and numerous land plants and the most commercially important group to humans. The major exception to the dominance of terrestrial ecosystems by flowering plants is the coniferous forest The temperate coniferous forest includes areas such as the Valdivian temperate rain forests of southwestern South America, the rain forests of New Zealand and Tasmania, northwest Europe , southern Japan, and the eastern Black Sea-Caspian Sea region of Turkey and Georgia to northern Iran.
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